He actually applies metallic industrial paints to his work using porcupine quills to get this rich detailed effect.

Lyon was the first to describe the Bornean white bearded gibbon ( Hylobates albibarbis ), the Gansu pika ( Ochotona cansus ), the Sumatran porcupine ( Hystrix sumatrae ), the shadowy broad nosed bat ( Platyrrhinus umbratus ), and two species of slow loris from Borneo, Nycticebus bancanus and Nycticebus borneanus .